Publication list also available at .
AFM image analysis software (SmarTrace, AutoSmarTrace) is available for download from .

Tumbleweed: an artificial motor protein that walks along a DNA track. P. Nilsson, N.O. Robertson, N. Gustafsson, R.B. Davies, C.W. Liew, A. Lyons, R. Eichhorn, C.S. Niman, G.A. Blab, E.H.C. Bromley, A.E. Whitten, A.P. Duff, J.P. Beech, P. Jönsson, T. Böcking, B. Höcker, D.N. Woolfson, N.R. Forde, H. Linke, P.M.G. Curmi.

Atomic Force Microscopy Investigations of Collagen: From Single Molecules to Fibrils. J. Srocka*, A. Al-Shaer* and N.R. Forde.

AGEing of Collagen: The Effects of Glycation on Collagen's Stability, Mechanics and Assembly. D. Sloseris and N.R. Forde. Matrix Biology, 135, 153-160 (2025)

Walking by design: how to build artificial molecular motors made of proteins. P.F. Nilsson, A. Zink, O.M.C. Laprévote, C.S. Korosec, N. Gustafsson, N.O. Robertson, N.R. Forde, R. Eichhorn, B. Höcker, P.M.G. Curmi and H. Linke Nano Futures, 8, 042501 (2024)

Motility of an autonomous protein-based artificial motor that operates via a burnt-bridge principle. [aka the Lawnmower!] C.S. Korosec, I. Unksov, P. Surendiran, R. Lyttleton, P.M.G. Curmi, C.N. Angstmann, R. Eichhorn, H. Linke and N.R. Forde. Nature Communications 15, 1511 (2024)

Fluence-dependent degradation of fibrillar type I collagen by 222 nm far-UVC radiation. A. Kowalewski and N.R. Forde. PLOS ONE 19, e0292298 (2024)
Line optical tweezers as controllable micromachines: techniques and emerging trends. Y. Shen, D.A. Weitz, N.R. Forde and M. Shayegan. Soft Matter 18, 5359-5365 (2022)

Through the Eyes of Creators: Observing Artificial Molecular Motors. I.N. Unksov, C.S. Korosec, P. Surendiran, D. Verardo, R. Lyttleton, N.R. Forde and H. Linke. ACS Nanosci. Au 2, 140–159 (2022)
Sequence-dependent mechanics of collagen reflect its structural and functional organization. A. Al-Shaer, A. Lyons, Y. Ishikawa, B.G. Hudson, S.P. Boudko and N.R. Forde. Biophysical Journal 120, 4013-4028 (2021)

AutoSmarTrace: Automated Chain Tracing and Flexibility Analysis of Biological Filaments. M. Schneider, A. Al-Shaer and N.R. Forde. Biophysical Journal 120, 2599-2608 (2021)

Multivalent Diffusive Transport. A. Kowalewski, N.R. Forde and C.S. Korosec. Journal of Physical Chemistry B 125, 6857–6863 (2021)

Substrate stiffness tunes the dynamics of polyvalent rolling motors. C.S. Korosec, L. Jindal, M. Schneider, I. Calderon de la Barca, M.J. Zuckermann, N.R. Forde and E. Emberly. Soft Matter 17, 1468-1479 (2021)

Optical tweezers approaches for probing multiscale protein mechanics and assembly. K. Lehmann, M. Shayegan, G.A. Blab and N.R. Forde. Frontiers in Molecular Biosciences 7, 577314 (2020)

Apparent superballistic dynamics in one-dimensional random walks with biased detachment. C.S. Korosec, D.A. Sivak and N.R. Forde. Physical Review Research 2, 033520 (2020)
Synthetic biology approaches to dissecting linear motor protein function: towards the design and synthesis of artificial autonomous protein walkers. H. Linke, B. Höcker, K. Furuta, N.R. Forde and P.M.G. Curmi. Biophysical Reviews, 12, 1041-1054 (2020)
Characterizing the Flexibility of the Body’s Building Block: Collagen. A. Al-Shaer, A. Lyons and N.R. Forde. Physics in Canada, 75, 14-17 (2019)

Mechanics and Structural Stability of the Collagen Triple Helix. M.W.H. Kirkness, K. Lehmann and N.R. Forde. Current Opinion in Chemical Biology 53, 98-105 (2019)
A chloride ring is an ancient evolutionary innovation mediating the assembly of the collagen IV scaffold of basement membranes. V. Pedchenko, R. Bauer, E.N. Pokidysheva, A. Al-Shaer, N.R. Forde, A.L. Fidler, B.G. Hudson and S.P. Boudko. Journal of Biological Chemistry 294, 7968-7981 (2019)
The Bar-Hinge Motor: A synthetic protein design exploiting conformational switching to achieve directional motility. L.S.R. Small, M.J. Zuckermann, R.B. Sessions, P.M.G. Curmi, H. Linke, N.R. Forde and E.H.C. Bromley. New Journal of Physics 21, 013002 (2019)

Modified Pluronic F127 surface for bioconjugation and blocking nonspecific adsorption of microspheres and biomacromolecules. M.H.W. Kirkness, C.S. Korosec and N.R. Forde. Langmuir 34, 13550-13557 (2018)

Environmentally controlled curvature of single collagen proteins. N. Rezaei*, A. Lyons* and N.R. Forde, Biophysical Journal 115, 1457-1469 (2018)

Dimensionality-dependent crossover in motility of polyvalent burnt-bridges ratchets. C.S. Korosec, M.J. Zuckermann and N.R. Forde, Physical Review E 98, 032114 (2018)
Fluorescence Correlation Spectroscopy (FCS) module for advanced undergraduate laboratories. N.R. Forde, D. Lee and J. Bechhoefer, 2018 Conference on Laboratory Instruction Beyond the First Year of College (BFY) Proceedings (2018)

Single-molecule assay for proteolytic susceptibility using centrifuge force microscopy: force-induced destabilization of collagen's triple helix. M.W.H. Kirkness and N.R. Forde. Biophysical Journal 114, 570–576 (2018)
Genetically modified human type II collagen for N- and C-terminal covalent tagging. A. Wieczorek, C.K. Chan, S. Kovacic, C. Li, T. Dierks and N.R. Forde. Canadian Journal of Chemistry 96, 204-211 (2018)

Effects of finite and discrete sampling and blur on microrheology experiments. V.E. Loosemore and N.R. Forde. Optics Expess 25, 31239-31252 (2017)
Engineering Nanoscale Biological Molecular Motors. C.S. Korosec and N.R. Forde. Physics in Canada 73, 78-81 (2017)
Gaining practical experience with Physics approaches to the micro- and nanoscale world of biology. N.R. Forde. Physics in Canada 73, 109-112 (2017)
Nanophysical Approaches to Biological Systems (foreword). J. Dutcher, N.R. Forde and S. Leslie. Physics in Canada 73, 59-60 (2017)
Construction of a chassis for a tripartite protein-based molecular motor. L.S.R. Small, M. Bruning, A.R. Thomson, A.L. Boyle, R. Davies, P.M.G. Curmi, N.R. Forde, H. Linke, D.N. Woolfson and E.H.C. Bromley. ACS Synthetic Biology 6, 1096-1102 (2017)

Intact telopeptides enhance interactions between collagens. M. Shayegan*, T. Altindal*, E. Kiefl and N.R. Forde. Biophysical Journal 111, 2404 (2016).
Development and characterization of a eukaryotic expression system for human type II procollagen. A. Wieczorek, N. Rezaei, C.K. Chan, C. Xu, P. Panwar, D. Brömme, E.F. Merschrod S. and N.R. Forde. BMC Biotechnology 15, 112 (2015).
Motor properties from persistence: a linear molecular walker lacking spatial and temporal asymmetry. M.J. Zuckermann, C.N. Angstmann, R. Schmitt, G.A. Blab, E.H.C. Bromley, N.R. Forde, H. Linke and P.M.G. Curmi. New Journal of Physics 17, 055017 (2015)
Design and construction of the lawnmower, an artificial burnt-bridges motor. S. Kovacic, L. Samii, P.M.G. Curmi, H. Linke, M.J. Zuckermann and N.R. Forde. IEEE Transactions on NanoBioscience 4, 305-312 (2015)
Probing Telopeptide-Induced Collagen-Collagen Interactions Using Optical-Tweezers-Based Microrheology. T. Altindal, M. Shayegan, E. Kiefl and N.R. Forde, in Optical Trapping Applications (Optical Society of America, Washington, DC, 2015), OtM3E.3
Fluidic Switching in Nanochannels for Control of Inchworm: A Synthetic Biomolecular Motor with a Power Stroke. C.S. Niman, M.J. Zuckermann, M. Balaz, J.O. Tegenfeldt, P.M.G. Curmi, N.R. Forde and H. Linke. Nanoscale 6, 15008-15019 (2014)
Construction and Characterization of Kilobasepair Densely Labeled Peptide-DNA. S. Kovacic, L. Samii, G. Lamour, H. Li, H. Linke, E.H.C. Bromley, D.N. Woolfson, P.M.G. Curmi and N.R. Forde. Biomacromolecules 15, 4065–4072 (2014)
Probing multiscale mechanics of collagen with optical tweezers. M. Shayegan, N. Rezaei, N.H. Lam, T. Altindal, A. Wieczorek and N.R. Forde. in Optical Trapping and Optical Micromanipulation X, edited by K. Dholakia and G.C. Spalding, Proceedings of SPIE 8810 88101P (2013)
Microrheological Characterization of Collagen Systems: From Molecular Solutions to Fibrillar Gels. M. Shayegan and N.R. Forde. PLOS ONE 8, e70590 (2013)
. C.S. Niman, J.P. Beech, J.O. Tegenfeldt, P.M.G. Curmi, D.N. Woolfson, N.R. Forde and H. Linke. Lab on a Chip 13, 2389-2396 (2013)
. A.L. Boyle, E.H.C. Bromley, G.J. Bartlett, R.B. Sessions, T.H. Sharp, C.L. Williams, P.M.G. Curmi, N.R. Forde, H. Linke and D.N. Woolfson. Journal of the American Chemical Society 134, 15457–15467 (2012)
. M. Shayegan and N.R. Forde. MRS Proceedings 1465, mrss12-1465-ss06-08 (2012)
. S. Kovacic*, L. Samii*, D. N. Woolfson, P.M.G. Curmi, H. Linke, N.R. Forde and G.A. Blab. Journal of Nanomaterials 2012, 109238 (2012)
. N.J. Kuwada, M.J. Zuckermann, E.H.C. Bromley, R.B. Sessions, P.M.G. Curmi, N.R. Forde, D.N. Woolfson and H. Linke, Physical Review E 84, 031922 (2011)
. A. Farré, M. Shayegan, C. López-Quesada, G.A. Blab, M. Montes-Usategui, N.R. Forde and E. MartÃn-Badosa, Optics Express 19, 21370-21384 (2011)
. L. Samii, G.A. Blab, E.H.C. Bromley, H. Linke, P.M.G. Curmi, M.J. Zuckermann and N.R. Forde, Physical Review E 84, 031111 (2011)
. N. Rezaei, B.P.B. Downing, A. Wieczorek, C.K.Y. Chan, R.L. Welch and N.R. Forde. in Photonics North 2011, edited by R. Kashyap, M. Têtu and R.N. Kleiman, Proceedings of SPIE 8007 80070K (2011)
. A. van der Horst and N.R. Forde. Optics Express 18, 7670-7677 (2010)
. A. Farré*, A. van der Horst*, G.A. Blab, B.P.B. Downing and N.R. Forde. Journal of Biophotonics 3, 224-233 (2010)
. L. Samii, H. Linke, M.J. Zuckermann and N.R. Forde, Physical Review E 81, 021106 (2010)
. E.H.C. Bromley, N.J. Kuwada, M.J. Zuckermann, R. Donadini, L. Samii, G.A. Blab, G.J. Gemmen, B.J. Lopez, P.M.G. Curmi, N.R. Forde, D.N. Woolfson and H. Linke, HFSP Journal 3, 204-212 (2009)
. B.P.D. Downing, A. van der Horst, M. Miao, F.W. Keeley and N.R. Forde, in Optical Trapping Applications (Optical Society of America, Washington, DC, 2009), OTuA3
. A. van der Horst, B.P.D. Downing and N.R. Forde, in Optical Trapping Applications (Optical Society of America, Washington, DC, 2009), OMB3
. A. van der Horst and N.R. Forde, Opt. Express 16, 20987-21003 (2008)
. Y.X. Mejia, H. Mao, N.R. Forde and C. Bustamante, J. Mol. Biol. 382, 628-637 (2008)
. Y. Deng, J. Bechhoefer and N.R. Forde, J. Opt. A: Pure Appl. Opt. 9, S256-S263 (2007) [special issue on optical trapping] Featured in the November/December 2007 issue of .
. Y. Deng, N.R. Forde and J. Bechhoefer, Phys. Rev. Lett. 98, 189082 (2007)
. C. Bustamante, Y.R. Chemla, N.R. Forde and D. Izhaky, Ann. Rev. Biochem. 73, 705-748 (2004) [authors listed alphabetically]
. N.R. Forde, D. Izhaky, G.R. Woodcock, G.J.L. Wuite and C. Bustamante, Proc. Natl. Acad. Sci. 99, 11682-11687 (2002)
Theses by group members
Chapin Korosec, Ph.D. in Physics, 2021
Michael Kirkness, Ph.D. in Molecular Biology and Biochemistry, 2019
Victoria Loosemore, M.Sc. in Physics, 2017
Naghmeh Rezaei, Ph.D. in Physics, 2016
Marjan Shayegan, Ph.D. in Chemistry, 2014
Laleh Samii, Ph.D. in Physics, 2013
Benjamin Downing, M.Sc. in Physics, 2010
Michelle Lee, M.Sc. in Physics, 2007
Yi Deng, M.Sc. in Physics, 2007
Philip Johnson, M.Sc. in Physics, 2007
Papers from Nancy's earlier life in Chemical Physics:
Characterization of nitrogen-containing radical products from the photodissociation of trimethylamine using photoionization detection. N.R. Forde, L.J. Butler, O. Sorkhabi, F. Qi, A. Suits and B. Ruscic, J. Chem. Phys. 113, 3088-3097 (2000)
Photodissociating Trimethylamine at 193 nm to Probe Dynamics at a Conical Intersection and to Calibrate Radical Detection Efficiency. N.R. Forde, M.L. Morton, S.L. Curry, S.J. Wrenn and L.J. Butler, J. Chem. Phys. 111, 4558-4568 (1999)
Electronic accessibility of dissociation channels in an amide: N,N-dimethylformamide photodissociation at 193 nm. N.R. Forde, L.J. Butler and S. A. Abrash, J. Chem. Phys. 110, 8954-8968 (1999)
Chemical Reaction Dynamics when the Born-Oppenheimer Approximation Fails: Understanding Which Changes in the Electronic Wavefunction might be Restricted. N.R. Forde, T.L. Myers and L.J. Butler, Faraday Discussions 108, 221-242 (1997)
Photodissociation of Acrylic Acid at 193 nm. D.C. Kitchen, N.R. Forde and L.J. Butler, J. Phys. Chem. A 101, 6603-6610 (1997)